Jugami Environmental Science

High School · Environmental Science

Ecosystems

How life, energy and matter connect.

On the rocky shore of Tatoosh Island, Washington, an ecologist named Robert T. Paine kept prying purple sea stars off the rocks and throwing them back into the sea. It sounds like a small thing to do. Within a year, the number of species living on those rocks had been cut in half.

How can losing one predator wreck a whole community? To answer that, you need to see an ecosystem the way ecologists do: as a web of living things and their surroundings, held together by flows of energy and cycles of matter. This lesson walks through that web piece by piece, then comes back to Paine's sea stars.

Drawing of a pond ecosystem. The sun shines on cattails, a tree, a frog on a lily pad and a heron in the shallows. Labels mark biotic parts, the living things, and abiotic parts: sunlight, water, air, rock and soil. Biotic: treefrogheroncattails Abiotic: sunlightwaterrocksoilair

On this page

  1. 1Biotic and abiotic factors
  2. 2Food chains and food webs
  3. 3Trophic levels and the 10% rule
  4. 4Nutrient cycles: carbon, nitrogen, water
  5. 5Biomes
  6. 6Succession
  7. 7Keystone species
  8. 8Human impacts
  9. 9Check your understanding
  10. 10Vocabulary

1. Biotic and abiotic factors

An ecosystem is a community of living things together with the nonliving environment around them, such as the weather, water and landscape, all interacting in one area. A pond is an ecosystem. So is a rotting log, a coral reef or the Amazon rainforest. Ecosystems have no fixed size; what makes something an ecosystem is that its parts interact.

Those parts come in two kinds:

Biotic = living

Biotic factors

Living things and things that were once alive: plants, animals, fungi, bacteria, and even fallen leaves and dead logs. In the pond in the picture above, the tree, cattails, frog and heron are biotic.

Abiotic = nonliving

Abiotic factors

The nonliving parts: sunlight, temperature, water, air, humidity, rocks and soil minerals. They set the rules for what can live in a place. Change the temperature or the rainfall, and the living community changes with it.

The two kinds constantly shape each other. Abiotic factors decide which organisms can survive, and organisms change their surroundings in return: tree roots break rock into soil, and the shade of a forest cools the ground beneath it.

2. Food chains and food webs

Every organism needs energy, and almost all of that energy starts as sunlight. Producers, also called autotrophs, capture it. Plants, algae and some bacteria use photosynthesis to turn light, carbon dioxide and water into sugar. (A few producers, mostly bacteria living in dark caves or at hydrothermal vents on the ocean floor, use chemical energy instead. They are called chemoautotrophs.) Everything else is a consumer that gets energy by eating.

A food chain is a single path of who eats whom, like grass → grasshopper → mouse → snake → hawk. Real nature is messier: mice eat more than one thing, and hawks hunt more than one animal. Linking all the food chains in an ecosystem gives a food web.

A grassland food web Arrows point from each food to the animal that eats it. Grass and clover are producers. Grasshoppers eat grass. Rabbits and mice eat grass and clover. Songbirds eat grasshoppers. Snakes eat mice. Red foxes eat mice and rabbits. Red-tailed hawks eat songbirds, snakes, mice and rabbits. Decomposers such as fungi, bacteria and earthworms break down dead matter from every level. Producers Primaryconsumers Secondaryconsumers Toppredator Grass Clover Grasshopper Rabbit Mouse Songbird Snake Red fox Red-tailed hawk Decomposersfungi, bacteria, worms Arrows point from food to eater, the way energy flows
A simplified grassland food web. Arrows point from the food to the eater, showing the direction energy moves. Dashed lines show a few examples of dead matter and waste going to the decomposers. In a real ecosystem, every level feeds them.

Two groups clean up after everyone else. Detritivores like earthworms and millipedes eat dead material, and decomposers, mainly fungi and bacteria, break it down into simple chemicals. Without them, dead matter would pile up and the nutrients locked inside it would never return to the soil.

Read the arrows the right way

The arrow in "grass → rabbit" does not mean "grass eats rabbit." It means energy moves from the grass into the rabbit. Follow the arrows and you are following the energy.

3. Trophic levels and the 10% rule

Each step in a food chain is a trophic level. Producers are the first level. Primary consumers (herbivores) eat producers. Secondary consumers eat herbivores, and tertiary consumers eat those. An animal at the very top with no natural predators of its own, like an orca or a hawk, is an apex predator. Many animals feed at more than one level; a fox that eats both berries and rabbits is a primary and a secondary consumer.

Here is the catch: energy does not pass up the chain completely. An animal uses most of the energy it eats just to stay alive, to move, grow, breathe and keep warm, and much of it ends up as heat. Some of what it eats is never digested at all. On average, only about 10% of the energy at one level is passed on to the next. This is the 10% rule.

An energy pyramid using the 10% rule Four stacked levels. Producers at the bottom hold 10,000 kilocalories. Primary consumers get about 1,000, secondary consumers about 100, and tertiary consumers at the top about 10. At each step about 90 percent of the energy is used for living or lost as heat. Tertiary10 kcal Secondary consumers100 kcal Primary consumers1,000 kcal Producers10,000 kcal ~90% lost ~90% lost ~90% lost Red arrows: at each step, most energy is used for living or lost as heat. Round numbers for illustration. Real ecosystems vary around the 10% average.
An energy pyramid. Each level holds roughly a tenth of the energy of the level below it.

Try it: the energy pyramid calculator

Start with the energy captured by producers and see how much is left at each level.

10% is the average used in the rule. Slide it to see what changes when an ecosystem is more or less efficient.

  • Level 5: 1 kcal
  • Level 4 (tertiary consumers): 10 kcal
  • Level 3 (secondary consumers): 100 kcal
  • Level 2 (primary consumers): 1,000 kcal
  • Level 1 (producers): 10,000 kcal

Level 5 gets 0.01% of the energy the producers captured.

That steep drop explains a lot. It is why food chains are short: after four to six energy transfers, there is not enough energy left to support another level. It is also why a meadow has far more rabbits than foxes.

How do we know? Measuring energy in a Florida spring

Ecologists tested these ideas at Silver Springs, Florida, by measuring the energy stored at each trophic level. Producers there captured 20,810 kilocalories per square meter per year (the gross primary productivity), but after using energy for their own respiration only 7,633 kcal/m²/yr was left as net primary productivity for the rest of the ecosystem. The transfer efficiency between the first two trophic levels was about 14.48%. That's a bit better than 10%, which is the point: "10%" is an average, not a law of physics. Real values vary from one ecosystem to the next, but they are always far below 100%.

Talk about it

Using the 10% rule, explain why eating plants directly can feed more people from the same farmland than feeding those plants to cattle and eating the beef.

4. Nutrient cycles: carbon, nitrogen and water

Energy makes a one-way trip through an ecosystem: in as sunlight, out as heat. Matter is different. The atoms that make up living things are not used up; they are recycled. The carbon, nitrogen and water in your body have been part of other living things before you, maybe ocean plankton or an ancient forest. These atoms move between living things, air, water and rock in biogeochemical cycles.

The carbon cycle

The carbon cycle Carbon dioxide in the atmosphere is taken in by plants during photosynthesis. Plants, animals and decomposers return it through respiration and decomposition. The ocean absorbs carbon dioxide from the air and releases some back. Over long times, carbon is buried as fossil fuels and limestone in the slow cycle. People release that buried carbon quickly by burning fossil fuels. CO₂ in the atmosphere Photosynthesis Respiration(plants, animals)and decomposition Burningfossil fuels Oceanexchange Ocean: dissolvedcarbon, plankton, shells Soil: dead matter and decomposers Slow cycle: fossil fuels and limestone, stored for millions of years
The carbon cycle. Black arrows are natural flows; red shows people moving buried carbon into the air by burning fossil fuels.

Carbon is the backbone of every living molecule. In the fast carbon cycle, plants and algae pull carbon dioxide (CO₂) out of the air during photosynthesis and build it into sugars. Animals eat the plants, and both plants and animals release CO₂ back to the air through respiration. When organisms die, decomposers break them down and release more.

The ocean is the giant in the cycle: it holds about 50 times more carbon than the atmosphere. CO₂ dissolves into seawater and comes back out, and some of the dissolved carbon combines with calcium to form calcium carbonate, the main ingredient of many sea shells. In the slow carbon cycle, carbon gets locked away for millions of years in limestone rock and in fossil fuels (coal, oil and natural gas), which formed from ancient buried life.

People have short-circuited the slow cycle. Burning fossil fuels, clearing forests and making cement release carbon that was stored underground for ages. Natural sinks, mainly the ocean and plants, soak up about half of the CO₂ humans emit each year. The rest stays in the air.

How do we know? The Mauna Loa record

Scientists have measured CO₂ in the air at the Mauna Loa Observatory in Hawaii since 1958, when it was about 315 parts per million (ppm). Before the Industrial Revolution it was about 280 ppm or less. In 2024 the global average reached a record 422.8 ppm, about 50% higher than pre-industrial levels, and that year's jump of 3.75 ppm was the largest on record. Because the ocean absorbs some of this CO₂, the average pH of surface ocean water has fallen from about 8.21 to 8.10, a change called ocean acidification.

The nitrogen cycle

The nitrogen cycle Nitrogen gas, N2, makes up most of the atmosphere but plants cannot use it. Nitrogen fixation by bacteria, including bacteria in the root nodules of legumes, and a small amount by lightning, turns it into ammonia and ammonium in the soil. Decomposers also release ammonium from waste and dead matter, called ammonification. Nitrifying bacteria turn ammonium into nitrite and then nitrate. Plants absorb nitrate and ammonium, called assimilation, and animals get nitrogen by eating plants. Denitrifying bacteria turn nitrate back into nitrogen gas. People add nitrogen with factory-made fertilizer, and runoff can carry extra nitrate into rivers. N₂ gas: about 78% of the air Lightning(small share) ← root nodules Fixation by bacteria Ammonium NH₄⁺ Nitrite NO₂⁻ Nitrate NO₃⁻ Nitrification (bacteria) Assimilation Ammonification (decomposers) Denitrification Factoryfertilizer
The nitrogen cycle. Bacteria do most of the work of moving nitrogen between the air, the soil and living things.

Living things need nitrogen to make proteins and DNA. There's no shortage of it: nitrogen gas (N₂) makes up about 78% of the atmosphere. The problem is that plants and animals can't use N₂ directly. Its two atoms are bonded too tightly. Turning it into a usable form takes special help, and most of that help comes from bacteria.

  1. Nitrogen fixation. Certain bacteria convert N₂ into ammonia (NH₃), which becomes ammonium (NH₄⁺) in soil. Some are free-living, such as cyanobacteria. Others, like Rhizobium, live in bumps called root nodules on legumes such as beans, peas and clover, trading nitrogen for the plant's sugar. Lightning fixes a small amount too.
  2. Nitrification. Other soil bacteria, such as Nitrosomonas, convert ammonium into nitrite (NO₂⁻), and then other bacteria turn nitrite into nitrate (NO₃⁻).
  3. Assimilation. Plants absorb ammonium and nitrate through their roots and build them into proteins. Animals get their nitrogen by eating plants or other animals.
  4. Ammonification. When organisms die or release waste, decomposers turn the nitrogen in it back into ammonium.
  5. Denitrification. Bacteria such as Pseudomonas and Clostridium, living where oxygen is scarce, convert nitrate back into N₂ gas, which returns to the air.

In 1909 chemists worked out the Haber-Bosch process, a way to fix nitrogen in factories. It made large-scale synthetic fertilizer possible. But fertilizer that washes off fields carries extra nitrogen (and phosphorus) into rivers, lakes and coastal water, which leads to the dead zones described in Human impacts.

The water cycle

The water cycle The sun warms the ocean and water evaporates. Plants release water vapor through transpiration. Vapor condenses into clouds, which drop precipitation as rain or snow on the mountains. Water runs off over land into rivers that flow back to the ocean, and some soaks into the ground by infiltration to become groundwater, which also slowly flows toward the sea. EvaporationTranspirationPrecipitationCondensationRunoff →Ocean Infiltration ↓Groundwater flows slowly to the sea →
The water cycle, driven by energy from the sun and by gravity.

The sun's energy evaporates water from oceans, lakes and soil, and plants release water vapor from their leaves through transpiration (together these are called evapotranspiration). The vapor cools and condenses into clouds, then falls as precipitation. Gravity takes over: water runs off the land into streams and rivers (runoff), or soaks into the ground (infiltration) to become groundwater.

About 96.5% of all Earth's water is salty ocean water. Of the small share that is fresh, more than 68% is frozen in ice caps and glaciers and about 30% is underground. People change the cycle too: pumping groundwater, building dams, paving land so rain runs off instead of soaking in, and warming the climate.

5. Biomes

A biome is a large region with a similar climate and similar kinds of plants and animals, wherever on Earth it appears. The two abiotic factors that matter most in defining land biomes are temperature and precipitation. Biologists usually recognize eight major terrestrial biomes. There are aquatic biomes too, divided into ocean (marine) and freshwater.

Eight major terrestrial biomes, their climate and typical plants
BiomeClimateTypical plants and features
Tropical wet forest (rainforest)Warm all year; 125–660 cm (50–200 in) of rain a yearBroad-leaved plants that shed and replace leaves throughout the year; very high productivity and species diversity
SavannaHot, with a long dry seasonGrasslands with scattered trees, in Africa, South America and northern Australia
Subtropical desertLess than 30 cm (12 in) of rain a year, sometimes as little as 2 cmVery dry; in some years evaporation is greater than precipitation. Daytime soil surface can top 60 °C (140 °F)
Chaparral (scrub forest)65–75 cm (25.6–29.5 in) of rain a year, mostly in winterDominated by shrubs; adapted to frequent fire. Found in California, around the Mediterranean and southern Australia
Temperate grasslandHot summers, cold winters; 25–75 cm (9.8–29.5 in) of precipitationGrasses with few trees except along streams; called prairies in North America and steppes in Eurasia
Temperate forestModerate temperatures; 75–150 cm (29.5–59 in) of rainDeciduous trees that drop their leaves each fall
Boreal forest (taiga)Long, cold winters; 40–100 cm (15.7–39 in) of precipitation, mostly as snowCone-bearing evergreens such as pine, spruce and fir
Arctic tundraVery cold; average winter temperature −34 °C (−29.2 °F); very little precipitationLow plants and lichens; permanently frozen ground called permafrost; a short growing season

Talk about it

The temperate grassland and the chaparral can get similar amounts of rain. Why might they still support such different plants? (Hint: look at when the rain falls.)

6. Ecological succession

Ecosystems are not frozen in place. After a disturbance, communities rebuild in a fairly predictable series of stages called ecological succession.

Stages of ecological succession Five panels from left to right. Bare rock or fresh lava. Lichens and mosses, the pioneers, begin to make thin soil. Grasses and wildflowers. Shrubs and young trees. A mature forest. Primary succession starts at the first panel with no soil. Secondary succession, after a fire or a cleared field, starts around the grasses stage because soil is already there. Bare rock or lava Lichens and mosses Grasses and wildflowers Shrubs and young trees Mature forest Time Primary starts hereSecondary often starts here
Succession from bare rock to forest. Each stage changes the soil, shade and moisture in ways that make room for the next.

Starts with no soil

Primary succession

Happens on brand-new land, like rock left by a retreating glacier or fresh lava in Hawaii. Hardy pioneer species move in first and slowly help build soil, which lets other plants follow.

Soil survives

Secondary succession

Happens after a disturbance that leaves soil behind, such as a wildfire or an abandoned farm field. Ash returns nutrients to the ground, and an oak and hickory forest cleared by fire can regrow far faster than a forest starting from bare rock.

How do we know? Watching life return to Mount St. Helens

On May 18, 1980, Mount St. Helens in Washington erupted and buried the land north of the volcano in hot pumice and ash, creating the Pumice Plain. Scientists have been studying its recovery ever since. Among the first pioneers they found, in 1981 and 1982, was the prairie lupine, a legume. Like other legumes, it hosts nitrogen-fixing bacteria, so it could grow in barren pumice and enrich the ground for the plants that came after. It is a living lab for primary succession.

7. Keystone species

Back to Tatoosh Island. Paine coined the term keystone species for a species that holds its ecosystem together, the way the keystone at the top of a stone arch keeps the whole arch from falling. Remove it, and the ecosystem changes dramatically.

Robert Paine's sea star experiment Two rocky shore patches side by side. Left, with the purple sea star present: a mix of mussels, barnacles, limpets, snails and seaweed. Right, after the sea stars were removed: mussels crowd out almost everything else. With sea stars: many species Sea stars removed: mussels take over Key: dark ovals = mussels, rings = barnacles, tan = limpets and snails,green = seaweed, purple star = Pisaster sea star (the keystone predator).
Paine's experiment, drawn as two rock patches. Without its main predator, one prey species took over.

How do we know? An experiment you can picture

The purple sea star Pisaster ochraceus is a major predator of mussels and barnacles on Tatoosh Island. When Paine removed the sea stars from a stretch of shore, mussels took over and crowded out other species, including the algae that fed sea snails, limpets and small clams. Within a year, the area's biodiversity was cut in half. Comparing those patches with untouched ones was the evidence that one predator can control a whole community.

A famous land example is the gray wolf in Yellowstone National Park. According to the National Park Service, at least 136 wolves were killed in the park between 1914 and 1926, and the last wolf pack was killed in 1926. Without wolves, elk populations exploded and overgrazed the plants along streams. From 1995 to 1997, 41 wild wolves from Canada and northwest Montana were released in the park. National Geographic reports that since then, elk populations have shrunk, willows have grown taller, and beaver and songbird populations have recovered. A chain of effects like this, running from a predator down through the food web, is called a trophic cascade.

Keystone species aren't always predators. Kelp is called a keystone host: kelp forests give shelter to sea otters and food to the animals otters eat, such as fish and sea urchins.

Talk about it

The National Park Service notes that wolf effects on elk are tangled up with other factors, such as other predators, how elk are managed outside the park, and weather like drought and harsh winters. What evidence would you want to collect to show that wolves, not something else, caused the willows to grow back?

8. Human impacts

People are part of ecosystems too, and we now shape almost every one on Earth. Each big impact connects to an idea from this lesson:

Carbon cycle

Climate change and ocean acidification

Burning fossil fuels and clearing land have pushed atmospheric CO₂ about 50% above pre-industrial levels, warming the planet and making the surface ocean more acidic.

Nitrogen cycle

Nutrient runoff and dead zones

Fertilizer and sewage feed explosive growth of algae and microbes in lakes and coastal water. As they grow and decompose, they use up the dissolved oxygen, and fish and shellfish die. More than 400 dead zones had been found worldwide as of 2008. One of the worst is in the Gulf of Mexico, fed by fertilizer runoff from the Mississippi River basin.

Food webs

Losing key species

Hunting, overfishing and habitat loss can remove keystone species and set off trophic cascades, as when Yellowstone lost its wolves.

Water cycle

Changing where water goes

Dams, pavement and heavy groundwater pumping change how water moves through the land, affecting streams, wetlands and the species that depend on them.

The same science points toward solutions. Restoring a missing species, as Yellowstone did, can help an ecosystem recover. Farmers can plant legumes to add nitrogen to soil naturally and use fertilizer more carefully so less runs off. And protecting forests and wetlands keeps the carbon and water cycles working in our favor.

Talk about it

Pick an ecosystem near your school, such as a park, a river or even a vacant lot. Name one biotic factor, one abiotic factor, one way people affect it, and one thing your community could do to help.

9. Check your understanding

Choose an answer for each question. With JavaScript on, you'll get feedback right away and your best score is saved on this device only. The full answer key is below the quiz.

1. A student lists things she found in a pond. Which one is an abiotic factor?

2. In a food web, what does an arrow from grass to rabbit show?

3. Producers in a meadow capture 20,000 kcal of energy. Using the 10% rule, about how much reaches the secondary consumers?

4. Why are food chains usually only a few links long?

5. Which process returns nitrogen gas (N₂) to the atmosphere?

6. Which process removes carbon dioxide from the atmosphere?

7. A wildfire burns a forest, but the soil survives and plants begin to grow back. What is this called?

8. What happened when Robert Paine removed sea stars from the shore of Tatoosh Island?

9. How does fertilizer runoff create a dead zone in coastal water?

10. Which biome has permafrost, permanently frozen ground?

Show the answer key
  1. C. The temperature of the water. Temperature is a nonliving condition. Frogs, algae and bacteria are all living, so they are biotic.
  2. A. Energy moves from the grass into the rabbit that eats it. Arrows show the direction energy flows: from the food to the eater.
  3. D. 200 kcal. 20,000 × 0.10 = 2,000 kcal for primary consumers, then 2,000 × 0.10 = 200 kcal for secondary consumers.
  4. B. So little energy is left after each transfer that it can't support another level. About 90% of the energy is used or lost as heat at each step, so after a few steps there isn't enough left.
  5. B. Denitrification. Denitrifying bacteria such as Pseudomonas turn nitrate back into N₂ gas. Fixation does the opposite.
  6. D. Photosynthesis. Plants and algae take in CO₂ during photosynthesis. The other three release CO₂.
  7. A. Secondary succession. Succession that starts with soil already in place is secondary. Primary succession starts on bare rock or lava with no soil.
  8. C. Mussels crowded out other species and biodiversity fell by half. Without their main predator, mussels took over and the number of species was cut in half within a year. That's what makes the sea star a keystone species.
  9. B. Extra nutrients fuel huge growth of algae and microbes, which use up the water's oxygen. Excess nitrogen and phosphorus cause overgrowth of algae and microbes, and their growth and decomposition deplete dissolved oxygen.
  10. C. Arctic tundra. The Arctic tundra has permafrost, very low precipitation and a growing season of only about 10 to 12 weeks.

10. Vocabulary

Abiotic factor
A nonliving part of an ecosystem, such as sunlight, temperature, water or soil minerals.
Apex predator
An animal at the top of a food web with no natural predators of its own.
Assimilation
The uptake of nitrogen compounds by plants, which build them into proteins and DNA.
Autotroph (producer)
An organism that makes its own food, usually by photosynthesis.
Biogeochemical cycle
The path an element or compound, such as carbon, nitrogen or water, takes between living things, air, water and rock.
Biome
A large region with a similar climate and similar kinds of plants and animals.
Biotic factor
A living or once-living part of an ecosystem.
Consumer (heterotroph)
An organism that gets energy by eating other organisms.
Decomposer
An organism, mainly fungi and bacteria, that breaks dead matter down into simple nutrients.
Denitrification
The conversion of nitrate back into nitrogen gas by bacteria.
Detritivore
An animal, such as an earthworm, that eats dead plant and animal material.
Ecosystem
A community of living things plus their nonliving environment, interacting in one area.
Eutrophication
Overgrowth of algae and microbes in water caused by excess nutrients, which can use up dissolved oxygen.
Food web
All the connected food chains in an ecosystem.
Keystone species
A species that has an outsized effect on its ecosystem, so removing it changes the whole community.
Nitrogen fixation
The conversion of nitrogen gas into ammonia or ammonium, mostly by bacteria.
Ocean acidification
The drop in ocean pH caused by seawater absorbing extra carbon dioxide.
Pioneer species
The first species to move into bare or disturbed land during succession.
Primary succession
Succession on new land with no soil, such as fresh lava or bare rock.
10% rule
The average pattern that only about 10% of the energy at one trophic level passes to the next.
Secondary succession
Succession after a disturbance that leaves the soil in place, such as a fire.
Transpiration
The release of water vapor from plant leaves.
Trophic cascade
A chain of effects through a food web caused by adding or removing a top predator.
Trophic level
A feeding step in a food chain, such as producer or primary consumer.

Sources

Facts and numbers in this lesson come from these sources. The diagrams are original drawings made for Jugami, based on the processes these sources describe.

  1. National Geographic Education, “Ecosystem.” education.nationalgeographic.org
  2. National Geographic Education, “Food Web.” education.nationalgeographic.org
  3. National Geographic Education, “Energy Flow and the 10% Rule.” education.nationalgeographic.org
  4. National Geographic Education, “The Role of Keystone Species in an Ecosystem.” education.nationalgeographic.org
  5. OpenStax, Biology 2e, 46.1 “Ecology of Ecosystems.” openstax.org
  6. OpenStax, Biology 2e, 46.2 “Energy Flow through Ecosystems” (Silver Springs data, transfer efficiency, chemoautotrophs). openstax.org
  7. OpenStax, Biology 2e, 46.3 “Biogeochemical Cycles” (nitrogen cycle, eutrophication and dead zones). openstax.org
  8. OpenStax, Biology 2e, 45.6 “Community Ecology” (primary and secondary succession). openstax.org
  9. OpenStax, Biology 2e, 44.3 “Terrestrial Biomes.” openstax.org
  10. NOAA Education, “Carbon cycle.” www.noaa.gov
  11. NOAA Climate.gov, Rebecca Lindsey, “Climate change: atmospheric carbon dioxide” (2025). www.climate.gov
  12. NASA Earth Observatory, “The Carbon Cycle.” earthobservatory.nasa.gov
  13. USGS Water Science School, “The Water Cycle.” www.usgs.gov
  14. USGS Water Science School, “How Much Water is There on Earth?” www.usgs.gov
  15. University of Florida IFAS Extension, “The Nitrogen Cycle” (biological and industrial nitrogen fixation, Haber-Bosch). ask.ifas.ufl.edu
  16. U.S. National Park Service, Yellowstone, “History of Wolf Management.” www.nps.gov
  17. U.S. National Park Service, Yellowstone, “Wolf Ecology.” www.nps.gov
  18. USGS, “Ecology of Mount St. Helens National Monument.” www.usgs.gov
  19. John G. Bishop and others, USDA Forest Service Pacific Northwest Research Station, research on prairie lupine on the Pumice Plain of Mount St. Helens (2005). www.fs.usda.gov